The first of the Punic Wars between Carthage and Rome broke out in 264 BC. It took place entirely on the island of Sicily, and arose when the Carthagininan city of Messana revolted against Carthage. The Romans intervened against the Carthagininan response, and responded in turn by beseiging many of their other Sicilian colonies. Carthage then attempted to raise the seiges with its navy, only to have it defeated, leaving them in an untenable position. in 241 BC, the city-states signed a treaty in which the Romans kept Sicily, and the Carthagininans paid war damages (both of which they could easily afford). In 238, however, Carthage's mercenary troops were acting rebellious, and Rome took advantage by seizing the island of Corsica (a move that even Roman historians remember as rash and unethical). Carthage became extremely angry, and sent the general Hamilcar (Father of Hannibal) and his son-in-law Hasdrubal to Spain to found colonies and build up a military presence. This escalation is what eventually led to the Second Punic War.